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Transition of Sporozoites into Liver
Stage-Like Forms Is Regulated by the RNA Binding Protein Pumilio


Many eukaryotic developmental and cell fate decisions that are effected

post-transcriptionally involve RNA binding proteins as regulators of translation

of key mRNAs. In malaria parasites (Plasmodium spp.), the

development of round, non-motile and replicating exo-erythrocytic liver stage

forms from slender, motile and cell-cycle arrested sporozoites is believed to

depend on environmental changes experienced during the transmission of the

parasite from the mosquito vector to the vertebrate host. Here we identify a

Plasmodium member of the RNA binding protein family PUF as

a key regulator of this transformation. In the absence of Pumilio-2 (Puf2)

sporozoites initiate EEF development inside mosquito salivary glands

independently of the normal transmission-associated environmental cues.

Puf2- sporozoites exhibit genome-wide transcriptional

changes that result in loss of gliding motility, cell traversal ability and

reduction in infectivity, and, moreover, trigger metamorphosis typical of early

Plasmodium intra-hepatic development. These data

demonstrate that Puf2 is a key player in regulating sporozoite developmental

control, and imply that transformation of salivary gland-resident sporozoites

into liver stage-like parasites is regulated by a post-transcriptional

mechanism.


Vyšlo v časopise: Transition of Sporozoites into Liver Stage-Like Forms Is Regulated by the RNA Binding Protein Pumilio. PLoS Pathog 7(5): e32767. doi:10.1371/journal.ppat.1002046
Kategorie: Research Article
prolekare.web.journal.doi_sk: https://doi.org/10.1371/journal.ppat.1002046

Souhrn

Many eukaryotic developmental and cell fate decisions that are effected

post-transcriptionally involve RNA binding proteins as regulators of translation

of key mRNAs. In malaria parasites (Plasmodium spp.), the

development of round, non-motile and replicating exo-erythrocytic liver stage

forms from slender, motile and cell-cycle arrested sporozoites is believed to

depend on environmental changes experienced during the transmission of the

parasite from the mosquito vector to the vertebrate host. Here we identify a

Plasmodium member of the RNA binding protein family PUF as

a key regulator of this transformation. In the absence of Pumilio-2 (Puf2)

sporozoites initiate EEF development inside mosquito salivary glands

independently of the normal transmission-associated environmental cues.

Puf2- sporozoites exhibit genome-wide transcriptional

changes that result in loss of gliding motility, cell traversal ability and

reduction in infectivity, and, moreover, trigger metamorphosis typical of early

Plasmodium intra-hepatic development. These data

demonstrate that Puf2 is a key player in regulating sporozoite developmental

control, and imply that transformation of salivary gland-resident sporozoites

into liver stage-like parasites is regulated by a post-transcriptional

mechanism.


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